Gephyromantis ceratophrys Ranomafana Grainy Frog
This species was split from Gephyromantis asper by Miguel Vences and a bunch of other folks in 2017.
Ranomafana National Park, Fianarantsoa province, MadagascarApril 28, 2007
Ranomafana Grainy Frog (Gephyromantis ceratophrys)
This is a small floor-dwelling species, very wiggly and jumpy. I chased it around for awhile before it stopped for a brief photo session.

Here is a list of all the reptiles and frogs I saw on this trip to Madagascar.

Online references:
Printed references: